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anthro arms_tied arthropod_webbing bent_over bodily_fluids bodysuit bound canid canine canine_genitalia canine_penis clothing dominant dominant_female dragon duo facial_horn fellatio female fur furrification genital_fluids genitals gwen_stacy hi_res horn knot licking male male/female mammal marvel nose_horn oral penile penis penis_lick pussy_juice pussy_juice_through_clothing sex skinsuit spider-gwen spider-man_(character) spider-man_(series) spider-man_2099 spider_web superhero_costume tight_clothing tongue tongue_out web_bondage wet wet_clothing white_body white_fur 2_penises 4_fingers absurd_res animal_genitalia animal_penis anus arm_spikes beard big_penis black_beard black_claws black_dragon black_hair black_horn blizzard_entertainment bodily_fluids brown_body brown_claws brown_horn brown_scales building butt claws cloaca cloacal_penis clubbed_tail countershade_chest countershade_crotch countershade_feet countershade_hands countershade_neck countershade_paws countershade_tail 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wrathion_the_black_prince_(warcraft) 6my animal_genitalia anthro anus buttplug chain claws clothing dragon duo facial_horn genitals gloves glowing glowing_eyes glowing_markings gynomorph handwear hi_res horn intersex intersex/male jewelry k.b.(kb2001uwu) kobold latex latex_clothing latex_gloves latex_stockings legwear licking licking_lips lizard lube male markings plug_(sex_toy) reptile scalie semi-anthro sex_toy sheath size_difference stockings thick_thighs tongue tongue_out tosho 2023 3_fingers 3_toes ambiguous_gender anthro armwear bandai_namco bandanna black_sclera blue_eyes blush bridal_gauntlets canid canine ceratopsian chest_tuft claws clothing cute_fangs digimon digimon_(species) dinosaur duo facial_horn fangs feet fingers fur gammamon horn japanese_text kerchief leg_markings mammal markings multicolored_body multicolored_fur nose_horn open_mouth open_smile ornithischian renamon reptile scalie sesaminfps simple_background smile tail tail_markings teeth text toes tuft two_tone_body two_tone_fur white_background white_body white_fur white_markings yellow_body yellow_eyes yellow_fur <3 <3_eyes 2023 4_arms 69_position absurd_res antennae_(anatomy) anthro anthro_on_anthro anthro_penetrated anthro_penetrating anthro_penetrating_anthro armwear arthropod barefoot bat beetle big_butt big_penis black_antennae black_body black_exoskeleton black_spots blue_armwear blue_clothing blue_handwear blue_legwear blush bodily_fluids breasts breath butt clothing comic cucujoid daryl_nimble duo erection exoskeleton eyes_closed facial_horn fangs feet felino fellatio female female_on_top female_penetrated fingerless_gloves foreskin genitals glans gloves green_body green_exoskeleton green_eyes hair handjob handwear hi_res horn insect ladybug legwear lying male male/female male_on_bottom male_penetrating male_penetrating_female mammal moan mostly_nude motion_lines multi_arm multi_limb multiple_angles nose_horn on_back on_bottom on_top oral panting penetration penile penis penis_grab phoebe_(felino) pink_inner_ear pink_nose ponytail purple_clothing purple_gloves purple_hair purple_handwear purple_penis red_glans saliva saliva_string sex sharp_teeth smile spots spotted_body spotted_exoskeleton teeth text thick_thighs white_armwear white_clothing anal anal_penetration animal_genitalia anthro backwards_baseball_cap backwards_hat baileyx balls baseball_cap beach big_balls big_dom_small_sub big_penis bottomwear bottomwear_down canid canine canis clothed clothing daxterdingo dingo dominant dragon duo eyes_closed facial_horn feet fur furred_dragon genitals hair hat headgear headwear hi_res horn huge_penis jewelry large_penetration male male/male mammal muscular muscular_anthro muscular_male necklace nose_horn nude open_mouth pants pants_down partially_clothed pawpads penetration penile penile_penetration penis penis_in_ass penis_tip red_hair seaside sheath shirt size_difference smaller_penetrated smile tank_top tongue tongue_out topwear trexlis_(character) yellow_body yellow_fur 2_penises anthro asymmetrical_breast_frottage balls belly big_breasts black_body blue_hair blue_nipples bodily_fluids breast_squish breasts breasts_frottage chameleon digital_media_(artwork) diphallism duo facial_horn fish foreskin genital_fluids genitals grey_belly gynomorph hair hi_res horizontal_diphallism horn intersex intersex/intersex jackie_(radvengence) jackson's_chameleon lizard marine multi_genitalia multi_penis nipple_piercing nipples non-mammal_breasts nose_horn nubbed_corona_glans nude partially_retracted_foreskin penis piercing purple_hair radvengence reptile scalie shadowed_eyes shark sharp_teeth smile squish teal_body teeth absurd_res anus balls big_anus big_balls big_butt big_penis bodily_fluids bovy_(character) butt cum cumshot darkgem dragon ear_tag ejaculation erection facial_horn facial_piercing feral genital_fluids genitals hi_res horn huge_penis looking_at_viewer looking_back looking_back_at_viewer lying male nose_horn nose_piercing nose_ring on_grass on_ground on_side orange_body orange_scales outside penis piercing presenting presenting_hindquarters rear_view ring_piercing scales scalie solo sweat sweaty_butt yellow_eyes 1:1 2023 anthro artist_name blush claws colored curled_tail digital_drawing_(artwork) digital_media_(artwork) dragon ears_down earwiggy english_text facial_horn feet feet_up fingers hi_res horn jambu_(wof) looking_at_another looking_pleasured lying male nose_horn on_back one_eye_closed open_mouth pink_body pink_scales pivoted_ears rainwing_(wof) scales scalie signature simple_background solo solo_focus solo_focused tail text tongue tongue_out white_background wings wings_of_fire wink 3_toes 4_fingers absurd_res anthro barefoot blush bodily_fluids chubby_male facial_horn feet fingers foreskin furrowed_brow genital_fluids genital_piercing genitals hi_res holding_penis horn kobold looking_at_viewer male multicolored_body nose_horn omorashi open_mouth peeing peeing_on_ground peeing_on_self penis penis_piercing piercing potty_dance prince_albert_piercing scales simple_background solo tears toes tucked_tail two_panel two_panel_image two_tone_body urine urine_pool veryloudopossum watersports wetting yellow_eyes yellow_urine anthro arwa_(radvengence) big_breasts bikini bovid breasts brown_body brown_fur caprine chameleon cleavage clothed clothing digital_media_(artwork) duo facial_horn female fur goat green_eyes gynomorph_(lore) hair hi_res horn jackie_(radvengence) lizard looking_at_viewer mammal nipple_outline non-mammal_breasts nose_horn purple_eyes purple_hair radvengence reptile scalie swimwear wavy_hair anthro arwa_(radvengence) big_breasts bovid breasts brown_body brown_fur caprine chameleon cleavage clothed clothing digital_media_(artwork) duo facial_horn female fur goat green_eyes gynomorph_(lore) hair hi_res horn jackie_(radvengence) lizard looking_at_viewer mammal nipple_outline non-mammal_breasts nose_horn nude purple_eyes purple_hair radvengence reptile scalie swimwear topless wavy_hair absurd_res anthro big_breasts big_nipples blue_hair breasts cybernetics cyborg diana_digma dragon facial_horn female green_body green_scales hair hi_res horn hybrid machine nipples nude scales simple_background smile solo wooody absurd_res anthro big_breasts big_nipples blue_hair breasts clothed clothing cybernetics cyborg diana_digma dragon facial_horn female green_body green_scales hair hi_res horn hybrid machine nipples nude scales simple_background smile solo swimwear wooody absurd_res anthro blue_hair breasts clothed clothing cybernetics cyborg diana_digma dragon facial_horn female green_body green_scales hair hi_res horn hybrid machine scales simple_background smile solo swimwear wooody absurd_res angels_with_scaly_wings anus bodily_fluids brown_body brown_scales bryce_(angels_with_scaly_wings) claws clothed clothing countershading dragon erection facial_horn feral genital_fluids genitals grin hi_res horn legwear looking_at_viewer maid_uniform male n1k0 nose_horn on_ground penis presenting presenting_anus presenting_penis scales scalie scar smile solo tail tapering_penis uniform 4_fingers 4_toes 7_horns abs absurd_res anal anal_penetration animal_genitalia anthro anthro_on_feral anthro_penetrated arwuld balls bestiality big_dom_small_sub big_penis black_body black_countershading black_fingers black_horn blue_body cloaca cloacal_penis countershade_face countershade_neck countershade_torso countershading dahurgthedragon dominant dominant_male dragon duo erection eyes_closed facial_horn feathers feet feral feral_penetrating feral_penetrating_anthro fingers forearms fur furred_dragon genitals grey_body grey_pawpads grey_plates grey_scales grey_scutes hair hand_on_leg hand_on_thigh hi_res holding_partner horn humanoid_pointy_ears larger_male leaning leaning_forward leaning_over looking_at_another looking_at_partner looking_down looming_over male male/male male_penetrated male_penetrating male_penetrating_male mane markings membrane_(anatomy) membranous_wings multi_horn muscular muscular_anthro muscular_male pawpads paws penetration penile_spines penis pink_body pink_feathers pink_mane pink_penis pink_tail_tuft plating prehensile_penis purple_penis raised_arm red_body red_feathers ridged_penis scaled_dragon scaled_forearms scaled_legs scaled_underbelly scales scutes sex simple_background sitting_on_another size_difference smile spread_legs spreading striped_body striped_feathers striped_markings striped_tail stripes tail tail_markings tail_tuft tenebrose thick_penis toes tuft unusual_anatomy unusual_genitalia unusual_penis white_body white_countershading white_horn wings yellow_eyes 4_fingers 4_toes 7_horns abs absurd_res anal anal_penetration animal_genitalia anthro anthro_on_feral anthro_penetrated arwuld balls bestiality big_dom_small_sub big_penis black_body black_countershading black_fingers black_horn blue_body bodily_fluids cloaca cloacal_penis countershade_face countershade_neck countershade_torso countershading cum cum_in_ass cum_inside cum_on_face dahurgthedragon dominant dominant_male dragon duo ejaculation erection eyes_closed facial_horn feathers feet feral feral_penetrating feral_penetrating_anthro fingers forearms fur furred_dragon genital_fluids genitals grey_body grey_pawpads grey_plates grey_scales grey_scutes hair hand_on_leg hand_on_thigh hi_res holding_partner horn humanoid_pointy_ears larger_male leaning leaning_forward leaning_over looking_at_another looking_at_partner looking_down looming_over male male/male male_penetrated male_penetrating male_penetrating_male mane markings membrane_(anatomy) membranous_wings multi_horn muscular muscular_anthro muscular_male pawpads paws penetration penile_spines penis pink_body pink_feathers pink_mane pink_penis pink_tail_tuft plating prehensile_penis purple_penis raised_arm red_body red_feathers ridged_penis scaled_dragon scaled_forearms scaled_legs scaled_underbelly scales scutes sex simple_background sitting_on_another size_difference smile spread_legs spreading striped_body striped_feathers striped_markings striped_tail stripes tail tail_markings tail_tuft tenebrose thick_penis toes tuft unusual_anatomy unusual_genitalia unusual_penis white_body white_countershading white_horn wings yellow_eyes 4_fingers 4_toes 7_horns absurd_res animal_genitalia arwuld big_penis catsy_(artist) claws clenched_teeth cloaca cloacal_penis dahurgthedragon dragon erection facial_horn feathers feet feral fingers forearms frustrated fur furred_dragon genitals grey_body grey_pawpads grey_plates grey_scales grey_scutes hair hand_on_stomach hi_res horn looking_at_genitalia looking_at_own_penis looking_at_penis looking_at_self lying male mane markings multi_horn narrowed_eyes nose_horn on_back one_eye_closed pawpads paws penile_spines penis pink_body pink_feathers pink_mane pink_penis pink_tail_tuft plating prehensile_penis red_body red_feathers retractable_claws ridged_penis scaled_dragon scaled_forearms scaled_legs scaled_underbelly scales scratches scratching scutes solo spread_legs spreading striped_body striped_feathers striped_markings striped_tail stripes tail tail_markings tail_tuft teeth toes tuft unusual_anatomy unusual_genitalia unusual_penis white_claws white_horn yellow_eyes absurd_res arwuld blep brown_body brown_fur brown_horn brown_wings countershade_chest countershade_fur countershade_neck countershading dahurgthedragon doggystyle dominant dominant_feral dominant_male dragon duo eyes_closed facial_horn feathers female female_penetrated feral forearms from_behind_position fur furred_dragon gloves_(marking) grey_pawpads hair hi_res horn leaning leaning_forward looking_at_another looking_at_partner lying male male/female male_penetrating male_penetrating_female mane mane_hair markings membrane_(anatomy) membranous_wings mounting on_front orange_mane pawpads paws penetration pink_body pink_feathers pink_pawpads pink_tongue rape_face red_body red_feathers scaled_forearms scaled_legs scaled_underbelly scales scalie sex simple_background small_horn smile spots striped_markings striped_tail stripes submissive submissive_female tail tail_markings tan_body tan_countershading terra tongue tongue_out western_dragon white_horn wings yellow_eyes absurd_res breasts caressing caressing_chin caressing_head claws clothing colored_nails convenient_censorship dashinvaine dragon duo facial_horn female feral frill_(anatomy) gold_(metal) gold_jewelry hi_res horn human jewelry legwear looking_at_another mammal mountain nails nipples nose_horn nude nude_human orange_eyes pattern_clothing pattern_legwear pattern_stockings pupils red_body red_claws red_clothing red_nails red_scales ribbons rock scales sitting_on_another slit_pupils spines stockings tattoo wyvern absurd_res ambiguous_gender arm_tattoo caressing_face clothing colored_nails dashinvaine dragon duo facial_horn female feral frill_(anatomy) gold_(metal) gold_jewelry hair hi_res horn human jewelry legwear lingerie mammal mountain nails nose_horn orange_eyes pattern_clothing pattern_legwear pattern_stockings pupils red_body red_clothing red_legwear red_nails red_scales red_stockings rock scales sharp_teeth sitting_on_another slit_pupils spines stockings straight_hair tattoo teeth wyvern anthro armor balls big_balls big_penis blush boots clothing dinosaur facial_horn footwear foreskin gauntlets genitals gloves handwear hi_res horn male nose_horn penis reptile scalie sega solo sonic_the_hedgehog_(series) spikes theropod thrash_the_t-rex tyrannosaurid tyrannosaurus tyrannosaurus_rex white_crest_(artist) 2023 5_toes anthro ball_tuft balls claws digital_drawing_(artwork) digital_media_(artwork) dragon drawller erection facial_horn feet genitals glans hi_res horn humanoid_genitalia humanoid_penis male muscular muscular_anthro muscular_male navel nipples nose_horn outside penis plant sitting solo tail teeth toe_claws toes tree tuft absurd_res accessory after_transformation all_fours anthro balls bert_(mmhorrors) blue_body blue_penis bodily_fluids chain chained cock_ring dragon drooling duo facial_horn feral gastropod genitals gold_(metal) hair hi_res horn hybrid jewelry knot long_hair male mmhorrors mollusk nipples nose_horn parasite penile_spines penis penis_accessory penis_jewelry restraints saliva saliva_string slug spiked_penis spikes spikes_(anatomy) tail tongue tongue_out white_hair worm 4_horns ace_(kamex) anal anal_penetration anthro berserker_retro black_claws black_markings blue_body blue_fur blue_hair blue_skin bodily_fluids bovid butt butt_markings caprine carrying carrying_another carrying_partner claws cum cum_in_ass cum_inside demon dipstick_ears dragon drooling duo ear_piercing ear_ring eye_glint facial_hair facial_horn facial_markings fangs fur genital_fluids glint glowing glowing_eyes goatee green_claws green_horn green_spikes hair head_markings horn hybrid jagged_mouth larger_male larger_penetrated long_hair male male/male mammal markings multi_horn multicolored_body multicolored_ears multicolored_fur multicolored_skin muscular muscular_anthro muscular_male penetration penile penile_penetration penis_in_ass piercing ring_(marking) ring_piercing saliva sharp_teeth size_difference skindentation smaller_male smile snout snout_markings spikes spikes_(anatomy) tail tail_hand teeth tiredpinkpanda two_tone_body two_tone_fur two_tone_skin unusual_anatomy unusual_tail white_body white_fur white_hair white_horn white_tuft 2023 anthro beak belt blue_clothing bone_frill boots brown_body brown_boots brown_clothing brown_footwear brown_scales ceratopsian clothed clothing collared_shirt curved_sword digital_drawing_(artwork) digital_media_(artwork) dinosaur doublet drawing dyna_soar facial_horn fantasy flintlock flintlock_rifle footwear frill_(anatomy) gun hand_on_hip hat head_crest head_frill headgear headwear hi_res holding_gun holding_object holding_rifle holding_weapon horn long_hat male melee_weapon mercenary nose_horn orange_background ornithischian phrygian_cap puffy_sleeves ranged_weapon red_clothing reptile saurian_(disambiguation) scales scalie sheathed_weapon signature simple_background sketch sketchy solo standing sword tail topwear vest weapon white_clothing 16:9 2023 aliasing ambiguous_gender anthro argon_vile autograph dialogue english_text eyebrows facial_horn footprint generation_1_pokemon grey_background grey_body grey_skin horn male_(lore) monster_mind nintendo nose_horn open_mouth pawprint pokemon pokemon_(species) red_eyes rhydon simple_background solo teeth text tongue upload_policy_2.7 widescreen anthro big_breasts black_hair breasts brown_hair canid canine diana_digma dragon ewgengster_(artist) facial_horn female fish forest fox fur genitals green_body green_scales group hair hi_res horn hybrid iris_(zepompom) lyra_(zepompom) male male/female mammal marine nipples nude open_mouth plant pussy scales sex shark smile tail teeth tree wings absurd_res animal_genitalia animal_penis anthro big_penis black_hair blue_penis bodily_fluids canine_genitalia canine_penis cum cum_on_face cum_on_own_face cum_on_penis cum_on_self cumshot dragon ejaculation erection facial_horn fur furred_dragon genital_fluids genitals grey_body hair hi_res horn knot male masturbation membrane_(anatomy) membranous_wings nose_horn nova_(novatehderg) penile penile_masturbation penis solo symbolicproto wings 2023 5_fingers anthro athletic athletic_anthro athletic_female athletic_wear blue_eyes bodily_fluids bodysuit breasts bulge cetana claws closed_smile clothed clothing collarbone crotch_shot dialogue digital_drawing_(artwork) digital_media_(artwork) dominant dominant_female english_text facial_horn fan_character female fin fingers fish grey_body grey_scales hair hi_res horn logo looking_at_viewer looking_down marik_azemus34 marine markings mouth_closed multicolored_body multicolored_scales muscular muscular_anthro muscular_female navel non-mammal_breasts nose_horn orange_background partially_clothed patreon purple_background scales shaded shark shark_tail short_hair simple_background simple_shading skimpy skinsuit skintight_clothing skintight_suit smile solo sunset sweat tattoo teal_eyes text tight_clothing tight_fit tomboy tribal tribal_markings tribal_tattoo two_tone_body two_tone_scales url water watermark wet white_body white_hair white_scales yellow_background 16:9 2023 2_fingers 3_fingers abra ambiguous_gender anthro argon_vile arthropod biped black_body black_eyes black_fur blob_creature blue_background blue_body blue_fur blue_skin bottomwear brown_body brown_bottomwear brown_clothing brown_fur brown_shorts brown_skin buizel canid claws clothed clothing cute_fangs dated dipstick_tail elemental_creature english_text eye_markings eyes_closed facial_horn facial_markings finger_claws fingers floating fur generation_1_pokemon generation_2_pokemon generation_3_pokemon generation_4_pokemon gesture green_body green_fur green_scales grey_body grey_skin grimer group grovyle hand_on_shoulder head_markings heracross horn insect kecleon light_body light_fur lizard lucario magnet magnezone mammal markings mask_(marking) metal_creature mineral_fauna monster_mind mustelid necktie nintendo nose_horn nude one_eye_closed open_mouth open_smile orange_necktie pokemon pokemon_(species) purple_body red_body red_clothing red_eyes red_scales red_shirt red_topwear reptile rhydon sandslash scales scalie screw semi-anthro shirt shorts simple_background sleeveless_shirt smeargle smile standing swimming_trunks swimwear tail tail_markings tan_body tan_skin text tongue topless topwear torso_markings v_sign white_clothing white_shirt white_topwear widescreen yellow_body yellow_scales yellow_sclera yellow_skin anal anal_fingering anthro anus bestiality bodily_fluids butt cum cum_in_pussy cum_inside destabilizer disembodied_hand disembodied_penis duo erection facial_horn facial_piercing female feral fingering fisting genital_fluids genitals grey_body grey_skin hi_res horn human human_on_feral humanoid_genitalia humanoid_penis interspecies male male/female mammal nose_horn nose_piercing nose_ring penetration penis piercing puffy_anus pussy pussy_juice rear_view rhinoceros ring_piercing shannon_(buzzedminotaur) spread_pussy spreading standing teat_grab teats thumb_in_ass tongue tongue_out vaginal vaginal_fisting 16:9 2023 4_fingers 4_toes absurd_res all_fours animal_genitalia animal_penis anus ass_up balls black_pawpads blue_anus blue_clitoral_hood blue_clitoris blue_genital_slit blue_genitals blue_internal blue_mouth blue_pawpads blue_pussy bodily_fluids brown_body brown_fur butt canid canine cervix chin_horn claws clitoral_hood clitoris cloud cum cum_drip cum_in_pussy cum_in_uterus cum_inside cum_on_penis cum_while_penetrated cumshot deep_penetration detailed detailed_background digital_media_(artwork) dominant dominant_male dorsal_fin dragon dripping dripping_pussy duo ear_fins ear_frill ears_back ejaculation erection eyes_closed facial_horn facial_markings feet female female_on_bottom female_penetrated feral feral_on_feral feral_penetrated feral_penetrating feral_penetrating_feral feruda_(farstaria) fin fingers folded_wings fox fridaflame frill_(anatomy) fur genital_fluids genitals glowing glowing_markings grass grey_body grey_fur head_markings head_spines hi_res hindpaw horn impregnation internal internal_vaginal interspecies knot knotting leaking_cum light lighting looking_at_another looking_at_partner looking_pleasured male male/female male_on_top male_penetrating male_penetrating_female mammal markings membrane_(anatomy) membranous_wings moan narrowed_eyes nose_horn nostrills octo_the_kitsune on_bottom on_top oozing_cum open_mouth orange_body orange_fur orgasm_face outside outside_sex ovum pawpads paws penetration penile penile_penetration penis penis_in_pussy penis_shot pink_penis pivoted_ears plant plated_scales plates pubic_mound purple_body purple_dragon purple_scales pussy quadruped raised_tail runes scales scalie sex shaded size_difference sky slit snout spade_tail sperm_cell spikes spikes_(anatomy) spines spread_legs spread_pussy spreading submissive submissive_female tail tail_fin teeth toes urethra uterus vaginal vaginal_canal vaginal_knotting vaginal_penetration western_dragon white_body white_fur widescreen wing_claws wings yellow_body yellow_eyes yellow_scales 16:9 2023 4_fingers 4_toes absurd_res all_fours animal_genitalia animal_penis anus ass_up balls black_pawpads blue_anus blue_clitoral_hood blue_clitoris blue_genital_slit blue_genitals blue_mouth blue_pawpads blue_pussy bodily_fluids brown_body brown_fur butt canid canine chin_horn claws clitoral_hood clitoris cloud cum cum_drip cum_in_pussy cum_inside cum_on_penis cum_while_penetrated detailed detailed_background digital_media_(artwork) dominant dominant_male dorsal_fin dragon dripping dripping_pussy duo ear_fins ear_frill ears_back ejaculation erection eyes_closed facial_horn facial_markings feet female female_on_bottom female_penetrated feral feral_on_feral feral_penetrated feral_penetrating feral_penetrating_feral feruda_(farstaria) fin fingers folded_wings fox fridaflame frill_(anatomy) fur genital_fluids genitals glowing glowing_markings grass grey_body grey_fur head_markings head_spines hi_res hindpaw horn impregnation internal interspecies knot leaking_cum light lighting looking_at_another looking_at_partner looking_pleasured male male/female male_on_top male_penetrating male_penetrating_female mammal markings membrane_(anatomy) membranous_wings moan narrowed_eyes nose_horn nostrills octo_the_kitsune on_bottom on_top oozing_cum open_mouth orange_body orange_fur orgasm_face outside outside_sex ovum pawpads paws penetration penile penile_penetration penis penis_in_pussy pink_penis pivoted_ears plant plated_scales plates pubic_mound purple_body purple_dragon purple_scales pussy quadruped raised_tail runes scales scalie sex shaded size_difference sky slit snout spade_tail sperm_cell spikes spikes_(anatomy) spines 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